Leading an expedition in Germany, Kougami unseals the resting place of one of the alchemists that created the Core Medals in an effort to retrieve a set of lost Medals. However, upon the seal being undone, millions of Cell Medals erupt into a tower while creating a magical barrier that causes parts of Japan to flip over. As the alchemist reveals her plans to become a new OOO, Eiji and his friends are thrown into the past and must find their way back home and stop the alchemist before she destroys the world!
as Eiji Hino / Kamen Rider OOO
as Shingo Izumi / Ankh
as Hina Izumi
as Akira Date / Kamen Rider Birth
as Shintarou Gotou
as Erika Satonaka
as Kousei Kougami
as Chiyoko Shiraishi
as Kiyoto Maki
as Uva
as Kazari
as Mezool
as Gamel
as Yoshimune Tokugawa
as Shun Wakaba
as Satsuki Wakaba / Gara
as Bell
as Punk
as Gentaro Kisaragi / Kamen Rider Fourze
as Yuuki Joujima
as Mezool (voice)
as Gara (voice)
as Nue Yummy (voice)
as Knights (voice)
as O Scanner (voice)
as Birth Driver (voice) / Birth Buster (voice) / Narrator (voice)
